1. Department of Plant Pathology, Kansas State University , Manhattan, KS 66506 , USA
2. Department of Plant Pathology and Crop Physiology, Louisiana State University Agricultural Center , Baton Rouge, LA 70803 , USA
3. Department of Horticulture and Natural Resources, Kansas State University , Manhattan, KS 66506 , USA
4. School of Biosciences, University of Exeter , Exeter, EX4 4QD , UK
5. The Sainsbury Laboratory, University of East Anglia, Norwich Research Park , Norwich NR4 7UH , UK