1. Department of Biology, Evolution and Optics of Nanostructures Group, University of Ghent , Ledeganckstraat 35, 9000 Gent , Belgium
2. Department of Recent Vertebrates, Royal Belgian Institute of Natural Sciences , Vautierstraat 29, 1050 Brussels , Belgium
3. Museum of Comparative Zoology, Harvard University , Cambridge, MA 02138 , USA
4. Naturalis Biodiversity Center , Darwinweg 2, 2333 CR Leiden , The Netherlands