1. Pennsylvania Cooperative Fish and Wildlife Research Unit, Pennsylvania State University, University Park, PA, USA
2. U.S. Geological Survey, Pennsylvania Cooperative Fish and Wildlife Research Unit, Pennsylvania State University, University Park, PA, USA
3. Department of Ecosystem Science and Management, Pennsylvania State University, University Park, PA, USA
4. Pennsylvania Game Commission, Harrisburg, PA, USA
5. Department of Conservation and Natural Resources, Bureau of Forestry, Harrisburg, PA, USA