1. Centre for Biodiversity Dynamics, Department of Biology, NTNU , Trondheim , Norway
2. UK Centre for Ecology and Hydrology , Bush Estate, Penicuik , United Kingdom
3. UK Centre for Ecology and Hydrology , Bush Estate, Penicuik, United Kingdom
4. Highland Ringing Group , Tain , United Kingdom
5. School of Biological Sciences, University of Aberdeen , Aberdeen , United Kingdom