1. Department of Systematics, Biodiversity and Evolution of Plants, Albrecht-von-Haller Institute of Plant Sciences, University of Goettingen, Untere Karspuele 2, 37073 Goettingen, Germany
2. Department of Botany, National Museum of Natural History, Smithsonian Institution, P.O. Box 37012, MRC 166, Washington, DC 20013-7012, USA
3. Pharmaceutical Biology, Julius-von-Sachs-Institute for Biosciences, University of Wuerzburg, Julius-von-Sachs-Platz 2, 97082 Wuerzburg, Germany
4. Department Botany and Molecular Evolution, Senckenberg Research Institute and Natural History Museum Frankfurt, Senckenberganlage 25, 60325 Frankfurt am Main, Germany