1. Sorbonne Université, CNRS UMR 7093, Laboratoire d'Océanographie de Villefranche-sur-Mer, Station Zoologique, Villefranche-sur-Mer 06230, France
2. Tokyo University of Marine Science and Technology, School of Marine Resources and Environment, 4-5-7 Konan, Minato-ku, Tokyo 108-8477, Japan