1. Key Laboratory of Molecular Epigenetics of the Ministry of Education (MOE), Northeast Normal University, Changchun, China
2. State Key Laboratory of Agrobiotechnology, Key Laboratory of Crop Heterosis and Utilization (MOE), Key Laboratory of Crop Genetic Improvement (Beijing Municipality), China Agricultural University, Beijing, China
3. College of Agronomy and Biotechnology, China Agricultural University, Haidian District, Beijing, China
4. Institut of Plant Science Paris-Saclay, Université Paris sud, Orsay, France