1. Institute for Plant Sciences of Montpellier (IPSiM), Univ Montpellier, CNRS, INRAE, Institut Agro , Montpellier 34060, France
2. CIRAD, UMR AGAP Institut , Montpellier 34398, France
3. Inria Project-team Virtual Plants, UMR AGAP, Univ Montpellier, INRIA, CIRAD, INRAE , Montpellier 34398, France
4. Inria and LIRMM, Univ Montpellier, CNRS , Montpellier 34095, France