1. State Key Laboratory of Cell Biology, Shanghai Institute of Biochemistry and Cell Biology, Center for Excellence in Molecular Cell Science, University of Chinese Academy of Sciences, Chinese Academy of Sciences , Shanghai 200031 , China
2. School of Life Science and Technology, ShanghaiTech University , Shanghai 201210 , China
3. Shanghai Advanced Research Institute, Chinese Academy of Sciences , Shanghai 201210 , China
4. Guangdong Institute of Intelligence Science and Technology , Hengqin, Zhuhai 519031 , China