1. Department of Mathematics and Statistics, University of Strathclyde , 16 Richmond St, Glasgow G1 1XQ , UK
2. School of Mathematical Sciences, Peking University , Beijing 100871 , China
3. Beijing International Center for Mathematical Research , Center for Quantitative Biology, Center for Machine Learning Research, , Beijing 100871 , China
4. Peking University , Center for Quantitative Biology, Center for Machine Learning Research, , Beijing 100871 , China