1. Haywood Community College , Clyde, NC 28721 , USA
2. USDA Forest Service (retired), Northern Research Station , Morgantown, WV 26506 , USA
3. Department of Biology, West Virginia University , Morgantown, WV 26506 , USA
4. Division of Forestry and Natural Resources, West Virginia University , Morgantown, WV 26506 , USA