1. Smithsonian Environmental Research Center, Smithsonian Institution, Edgewater, MD 21037, USA
2. School of Biological Sciences
3. Central Science Laboratory, University of Tasmania, Hobart, TAS 7001, Australia
4. Graduate School of Animal and Food Hygiene, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido 080-8555, Japan
5. Graduate School of Agriculture, Tohoku University, Sendai, Miyagi 981-8555, Japan