1. CBGP, CIRAD, INRAE, IRD, Institut Agro, Univ. Montpellier , Montpellier , France
2. School of Agricultural, Earth and Environmental Sciences, University of KwaZulu-Natal , Durban , South Africa
3. CBGP, INRAE, CIRAD, IRD, Montpellier Institut Agro, Univ. Montpellier , Montpellier , France