1. International Research Centre for Environmental Membrane Biology & Department of Horticulture, Foshan University , Foshan 528000 , China
2. Tasmanian Institute of Agriculture, University of Tasmania , Hobart, TAS 7001 , Australia
3. Australian Research Council Centre of Excellence for Plant Success in Nature and Agriculture, School of Natural Sciences, University of Tasmania , Hobart, TAS 7001 , Australia
4. School of Biological Science, University of Western Australia , Crawley, WA 6009 , Australia