1. Shandong University School of Software, , Jinan 250101, China
2. Joint SDU-NTU Centre for Artificial Intelligence Research (C-FAIR), Shandong University , Jinan 250101, China
3. Institute of Fundamental and Frontier Sciences, University of Electronic Science and Technology of China , Chengdu 610054, China
4. Human Genome Center, Institute of Medical Science, The University of Tokyo , Tokyo 108-8639, Japan