1. National Key Laboratory of Agricultural Microbiology, College of Veterinary Medicine, Huazhong Agricultural University , Wuhan 430070 , China
2. Hubei Animal Disease Prevention and Control Center , Wuhan 430070 , China
3. Cooperative Innovation Center of Sustainable Pig Production, College of Veterinary Medicine , Wuhan 430070 , China
4. International Research Center for Animal Disease (Ministry of Science & Technology of China), College of Veterinary Medicine , Wuhan 430070 , China
5. The HZAU-HVSEN Research Institute , Wuhan 430042 , China