1. Department of Biological Sciences, Carnegie Mellon University , Pittsburgh, PA, USA
2. Instituto de Biomedicina de Sevilla, Hospital Universitario Virgen del Rocío/CSIC/Universidad de Sevilla , Seville, Spain
3. Departamento de Genética, Universidad de Sevilla , Seville, Spain
4. State Key Laboratory of Membrane Biology, Peking-Tsinghua Joint Centre for Life Sciences, School of Life Sciences, Peking University , Beijing, China