1. Division of Ecology and Evolution, Research School of Biology, The Australian National University , Canberra, ACT , Australia
2. Liaoning Key Laboratory for Biological Invasions and Global Changes, College of Bioscience and Biotechnology, Shenyang Agricultural University , Shenyang, Liaoning Province 110866 , China
3. Evolutionary Biology Group, Faculty of Biology, Adam Mickiewicz University , Poznań , Poland
4. Institute of Ecology and Evolution, School of Biological Sciences, University of Edinburgh , Edinburgh EH9 3FL , UK