1. RIKEN Center for Sustainable Resource Science (CSRS) , Yokohama 230-0045 , Japan
2. Institute of Plant and Microbial Biology, Academia Sinica , Taipei 11529 , Taiwan
3. Molecular and Biological Agricultural Sciences Program, Taiwan International Graduate Program, Academia Sinica and National Chung Hsing University , Taipei 11529 , Taiwan
4. Graduate Institute of Biotechnology, National Chung Hsing University , Taichung 40227 , Taiwan
5. Department of Biological Sciences, Graduate School of Science, The University of Tokyo , Tokyo 113-8654 , Japan
6. Biotechnology Center, National Chung Hsing University , Taichung 40227 , Taiwan