1. Université Paris-Saclay, INRAE, AgroParisTech, Institut Jean-Pierre Bourgin (IJPB) , 78000, Versailles , France
2. Institute of Plant Biology and Biotechnology, University of Münster , Schlossplatz 8, D-48143 Münster , Germany
3. Laboratory of Plant Molecular Genetics, Graduate School of Agricultural and Life Science, The University of Tokyo , Tokyo, 113-8657 , Japan