1. Keio University School of Medicine Laboratory of Cell and Tissue Biology, , 3N7, 35 Shinanomachi, Shinjuku-ku, Tokyo 160-8582, Japan
2. Hokkaido University Department of Biochemistry, Faculty of Medicine and Graduate School of Medicine, , Hokkaido 060-8636, Japan
3. National Institute for Physiological Sciences, National Institutes of Natural Sciences , Aichi 444-8787, Japan