1. Department of Biology and the Ecology Center, Utah State University, 5305 Old Main Hill, Logan UT 84321 USA
2. Department of Biological Sciences, Florida Gulf Coast University, 10501 FGCU Blvd S, Fort Myers, FL 33965 USA
3. Biological Sciences Department, Dixie State University, 225 S. University Avenue, St. George, UT 84770 USA