1. ComEU Sorbonnes Universités, UMR 7221 CNRS/MNHN, Muséum National d’Historie Naturelle, 7 rue Cuvier, 75231 Paris Cedex 05, France
2. Department of Evolutionary Ecology, Leibniz Institute for Zoo and Wildlife Research, Alfred-Kowalke-Straße 17, 10315 Berlin, Germany
3. Department of Biology, University of Antwerp, Universiteitsplein 1, 2610 Antwerp, Belgium
4. Institute of Biodiversity, Animal Health and Comparative Medicine, University of Glasgow, Glasgow G12 8QQ, UK
5. Department of Wildlife Diseases, Leibniz Institute for Zoo and Wildlife Research, Alfred-Kowalke-Straße. 17, 10315 Berlin, Germany