1. University of California Department of Ecology, Evolution & Marine Biology, , Santa Barbara, Santa Barbara, CA 93106-9620, USA
2. The University of British Columbia Pacific Salmon Ecology and Conservation Laboratory, Department of Forest and Conservation Sciences, , Vancouver, BC V6T 1Z4, Canada
3. Carleton University Fish Ecology and Conservation Physiology Laboratory, Department of Biology, , Ottawa, ON K1S 5B6, Canada
4. University of Gothenburg Department of Biological and Environmental Sciences, , 41390 Gothenburg, Sweden
5. Simon Fraser University Fisheries and Oceans Canada, Science Branch, Cooperative Resource Management Institute, School of Resource and Environmental Management, , Burnaby, BC V5A 1S6, Canada