1. Swire Institute of Marine Science and School of Biological Sciences, The University of Hong Kong, Pokfulam, Hong Kong
2. Marine Climate Change Unit, Okinawa Institute of Science and Technology Graduate University, Tancha, Onna-son, Okinawa 904-0495, Japan
3. ARC Centre of Excellence for Coral Reef Studies, James Cook University, 1 James Cook Dr, Townsville, Queensland 4814, Australia
4. Section for Physiology and Cell Biology, Department of Biosciences, University of Oslo, Blindernveien 31, 0316 Oslo, Norway