1. Max Planck Institute for Evolutionary Biology, Department of Evolutionary Ecology, August- Thienemann- Str. 2, D-24306, Ploen, Germany,
2. Department of Life and Environmental Science, Faculty of Science and Technology, Talbot Campus, Poole, BH12 5BB, UK,
3. GEOMAR| Helmholtz Centre for Ocean Research, Department of Evolutionary Ecology of Marine Fishes, D-24105, Kiel, Germany,
4. Present address: School of Biological and Chemical Sciences, Queen Mary University of London, Mile End Road, London E1 4NS, UK, and
5. Max Planck Institute of Immunobiology and Epigenetics, Department of Developmental Immunology, Stuebeweg 51, D-79108 Freiburg, Germany