1. Department of Biological Sciences, Macquarie University , Wallumattagal Campus, Macquarie Park, Culloden Rd 205 BCR, NSW 2000 Sydney , Australia
2. Behavioral Biology, Groningen Institute for Evolutionary Life Sciences, University of Groningen , Linnaeusborg (1st floor), Nijenborgh 7, 9747 AG Groningen , The Netherlands