1. Donghu Experimental Station of Lake Ecosystems, State Key Laboratory of Freshwater Ecology and Biotechnology of China, Institute of Hydrobiology, Chinese Academy of Sciences , Wuhan 430072 , China
2. Organismal and Evolutionary Biology, University of Helsinki , Helsinki, PO Box 65, 00014 , Finland
3. University of the Chinese Academy of Sciences , Beijing 100049 , China
4. Key Laboratory of Poyang Lake Environment and Resource Utilization of Ministry of Education, Nanchang University , Nanchang 330031 , China