1. Department of Biology, Terrestrial Ecology Unit (TEREC), Ghent University, K.L. Ledeganckstraat, Gent, Belgium
2. Laboratory of Socioecology and Socioevolution, KU Leuven, Naamsestraat, Leuven, Belgium
3. Research Unit of Environmental and Evolutionary Biology, Namur Institute of Complex Systems, and Institute of Life, Earth, and the Environment, University of Namur, Rue de Bruxelles, Namur, Belgium