1. Department of Natural Resources and the Environment, University of New Hampshire, Durham, New Hampshire, United States
2. School for Environment and Sustainability, University of Michigan, Ann Arbor, Michigan, United States
3. Conservation Biology Research Group, School of Environmental and Life Sciences, University of Newcastle, Callaghan, New South Wales, Australia
4. Mammal Research Centre, University of Pretoria, Tshwane, South Africa, and with the Centre for African Conservation Ecology, Nelson Mandela University, Port Elizabeth, South Africa
5. Wildlife Conservation Research Unit, Department of Zoology, University of Oxford, The Recanati-Kaplan Centre, Tubney, United Kingdom