1. Research Hub for Coral Reef Ecosystem Functions, with the College of Science and Engineering, and with the ARC Center of Excellence for Coral Reef Studies, James Cook University , Townsville, Queensland , Australia
2. Department of Biodiversity, Conservation, and Attractions, Kensington , Western Australia , Australia
3. UWA Oceans Institute, University of Western Australia, Crawley , Western Australia , Australia
4. Australian Institute of Marine Science, Indian Ocean Marine Research Centre, Crawley , Western Australia , Australia
5. UWA Oceans Institute, School of Biological Sciences, University of Western Australia, Crawley , Western Australia , Australia