1. School of Ocean Sciences, Bangor University , Menai Bridge, Isle of Anglesey, LL59 5AB , UK
2. Alfred Wegener Institute, Helmholz-Zentrum für Polar- und Meeresforschung, Marine Station Helgoland , 27498, Helgoland , Germany
3. Department of Marine Science, Faculty of Science, Kuwait University , Al-Fintas , Kuwait