1. Institute of Marine Biology, National Taiwan Ocean University , Keelung 20224 , Taiwan
2. Center of Excellence for the Oceans, National Taiwan Ocean University , Keelung 20224 , Taiwan
3. Graduate School of Integrated Sciences for Life, Hiroshima University , Higashi-Hiroshima, 739-8528 Hiroshima , Japan
4. School of Applied Biology, Hiroshima University , Higashi-Hiroshima, Hiroshima 739 - 8528 , Japan
5. Department of Biological Resources, National Chiayi University , Chiayi, 600355 , Taiwan