1. Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, 113-8657 Japan
2. Faculty of Science, Hokkaido University, Sapporo, 060-0810 Japan
3. Graduate School of Life Science, Hokkaido University, Sapporo, 060-0810 Japan
4. Department of Biological Chemistry, College of Bioscience and Biotechnology, Chubu University, Kasugai, 487-8501 Japan
5. Graduate School of Environmental Science, Hokkaido University, Sapporo, 060-0810 Japan
6. Graduate School of Agriculture, Hokkaido University, Sapporo, 060-8589 Japan