1. Universidade Federal do Pampa, Campus São Gabriel, Avenida Antônio Trilha, Centro, Brasil
2. Biological Laboratory, Faculty of Education, Nagasaki University, Nagasaki, Japan
3. Muséum national d’Histoire naturelle, Paris, France
4. Laboratório de Biologia Comparada e Abelhas, Faculdade de Filosofia Ciências e Letras de Ribeirão Preto, Universidade de São Paulo, Av. Bandeirantes, São Paulo, Brazil
5. Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, China
6. Institut de Recherche pour le Developpement (IRD, UR072 BP 1857), Yaoundé, Cameroon
7. Muséum national d’Histoire naturelle, MECADEV - UMR 7179 MNHN/CNRS, Paris, France