1. Department of Integrative Biology, University of California Berkeley , Berkeley, CA 94720, USA
2. Department of Biology, University of Victoria , Victoria, BC V8P 5C2, Canada
3. National Marine Fisheries Service, Alaska Fisheries Science Center, National Oceanographic and Atmospheric Administration , Auke Bay Laboratories, Juneau, AK 99801, USA
4. Center for Computational Biology, University of California Berkeley , Berkeley, CA 94720, USA