1. Department of Paleontology, University of Zurich , 8006 Zurich , Switzerland
2. Senckenberg Centre for Human Evolution and Palaeoenvironment an der Universität Tübingen , Tübingen , Germany
3. Fachbereich Geowissenshcaten dr Universität Tübingen , 72074 Tübingen , Germany
4. School of Biological Sciences, The University of Western Australia , Perth, WA 6009 , Australia
5. The UWA Oceans Institute, The University of Western Australia , Perth, WA 6009 , Australia