1. Radboud university medical center, Radboud Institute for Health Sciences, Department of Medical Microbiology , Nijmegen , Netherlands
2. Medical Research Council Unit The Gambia at the London School of Hygiene and Tropical Medicine , Banjul , The Gambia
3. LPHI, MIVEGEC, Université de Montpellier, CNRS , INSERM, Montpellier , France
4. Department of Infection Biology, London School of Hygiene and Tropical Medicine , London , United Kingdom