1. School of Biological Sciences, University of Bristol, Bristol Life Sciences Building, 24 Tyndall Avenue, Bristol, BS8 1TQ, UK
2. Institute of Systems, Molecular and Integrative Biology, Faculty of Health and Life Sciences, University of Liverpool, BioSciences Building, Crown Street, Liverpool, L69 7ZB, UK
3. Earlham Institute, Norwich Research Park, Norwich NR4 7UZ, UK
4. John Innes Centre, Norwich Research Park, Norwich NR4 7UH, UK
5. Department of Bioinformatics, Wissenschaftszentrum Weihenstephan, Technical University of Munich, 85354 Freising, Germany