1. Laboratory of Microbiology, Wageningen University and Research, Stippeneng 4, 6708 WE Wageningen, The Netherlands
2. Department of Biotechnology, Delft University of Technology, Van der Maasweg 9, 2629 HZ Delft, The Netherlands
3. DSM Biotechnology Center, Alexander Fleminglaan 1, 2613 AX Delft, The Netherlands