1. Oxford Centre for Computational Neuroscience, Oxford, UK
2. Department of Computer Science, University of Warwick, Coventry CV4 7AL, UK
3. Institute of Science and Technology for Brain-inspired Intelligence, Fudan University, Shanghai 200433, China
4. Key Laboratory of Computational Neuroscience and Brain-Inspired Intelligence, Fudan University, Ministry of Education, Shanghai 200433, China
5. School of Mathematical Sciences, School of Life Science and the Collaborative Innovation Center for Brain Science, Fudan University, Shanghai 200433, China