1. Departments of Medicine, Brigham and Women's Hospital and Harvard Medical School , Boston , MA 02115 , USA
2. Program in Virology, Harvard Medical School , Boston , MA 02115, USA
3. Instituto de Medicina Molecular, Faculdade de Medicina, Universidade de Lisboa , Avenida Professor Egas Moniz, 1649-028 Lisboa, Portugal
4. Universidade Católica Portuguesa, Católica Medical School , Católica Biomedical Research, Palma de Cima, 1649-023 Lisboa, Portugal
5. Broad Institute of Harvard University and Massachusetts Institute of Technology , Cambridge , MA 02142, USA