1. Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA
2. Behavior, Ecology, Evolution, and Systematics Program, University of Maryland, College Park, MD 20742, USA
3. Department of Biology, George Mason University, Manassas, VA 22030, USA
4. Center for Conservation Genomics, Smithsonian Conservation Biology Institute, Washington, DC 20008, USA