1. San Diego Natural History Museum, San Diego, CA 92101, USA
2. Department of Ecology, Evolution and Marine Biology, University of California, Santa Barbara, CA 93106, USA
3. Oak Creek Lab of Biology, Department of Fisheries, Wildlife and Conservation Sciences, Oregon State University, Corvallis, OR 97331, USA
4. Department of Land Resources and Environmental Sciences, Montana State University, Bozeman, MT 59717-3120, USA