1. Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, Harvard University, 26 Oxford Street, Cambridge MA 02138, USA
2. Smithsonian Tropical Research Institute, Panama City, Panama
3. Molecular Invertebrate Systematics and Ecology, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan
4. Tropical Biosphere Research Center, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan