1. Natural History Museum, University of Oslo , P.O. Box 1172, Blindern, 0318 Oslo , Norway
2. Centre of Molecular Biodiversity Research, Zoological Research Museum Alexander Koenig Bonn 53113 , Germany
3. FB05 Biology/Chemistry; University of Osnabrück , Osnabrück 49069 , Germany
4. Department of Natural History, University Museum of Bergen, University of Bergen , P.O. Box 7800, 5020 Bergen , Norway
5. Zoology Department, Faculty of Science, Damietta University , New Damietta, Central zone, 34517 , Egypt