1. Institute of Evolutionary Biology, University of Edinburgh, Ashworth Laboratories, King’s Buildings, Edinburgh, EH9 3FL, UK
2. Department of Computer Science, University of Auckland, 303/38 Princes St, Auckland, 1010, NZ
3. Centre for Computational Evolution, University of Auckland, 303/38 Princes St, Auckland, 1010, NZ
4. Department of Microbiology and Immunology, Rega Institute, KU Leuven - University of Leuven, Herestraat 49, 3000 Leuven, Belgium
5. Department of Human Genetics, University of California, Los Angeles, 695 Charles E. Young Dr., Los Angeles, CA 90095, USA
6. Department of Biostatistics, University of California, Los Angeles, 650 Charles E. Young Dr., Los Angeles, CA 90095, USA