1. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Center for Ocean Mega-Science, Institute of Oceanology, Chinese Academy of Sciences , Qingdao 266071 , China
2. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology , Qingdao 266237 , China
3. Key Laboratory of Breeding Biotechnology and Sustainable Aquaculture, Chinese Academy of Sciences , Qingdao 266071 , China
4. College of Earth Sciences, University of Chinese Academy of Sciences , Beijing 100049 , China