1. Department of Plant, Soil and Microbial Sciences, Michigan State University, East Lansing, MI 48824, USA
2. W.K. Kellogg Biological Station, Department of Integrative Biology, Michigan State University, Hickory Corners, MI 49060, USA
3. Department of Plant Pathology, Washington State University, Pullman, WA 99164, USA
4. Department of Crop and Soil Sciences, Washington State University, Pullman, WA 99164, USA