1. National Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences , Qingdao 266071, Shandong , China
2. Laboratory for Marine Fisheries Science and Food Production Processes, Marine Life research center, Laoshan Laboratory , Qingdao 266237, Shandong , China
3. College of Fisheries and Life Science, Dalian Ocean University , Dalian, Liaoning , China
4. Dalian Tianzheng Industry Co. , Ltd., Dalian, Liaoning , China